% replaces any number of letters (la% - lake, lamp)_ replaces one letter (ca_ - car, cat)
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The Crossword Solver found 10 answers for The New York Times crossword clue be in harmony 4 letters. The answers have between 3 and 7 letters.

AnswerClue
ALTOvocal part that often provides harmony to a soprano melody
MESHwork harmoniously
JIBEbe in harmony
DISCORDlack of harmony
BEEGEES“Britain’s first family of harmony,” per Brian Wilson
TONALlike music that uses conventional keys and harmony
ATTUNEDmade harmonious
THA“___ Crossroads” (Bone Thugs-N-Harmony hit that got a Grammy)
ONKEYharmonious, in a way
ATTUNEmake harmonious

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